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10434 Ancillary Reactions to the Pathway of fi -Oxidation P-Oxidation of saturated fatty acids generates an unsaturated intermediate with a A2* --tr-ans double bond, which is then hydrated by enoyl-CoA hydratase The metabolism of unsaturated and polyunsaturated fatty acids requires additional enzymes to act on preexisting cis double bonds As illustrated in Figure 10-7, the oxidation of linoleic acid (c9,c12-18:2) proceeds in essentially the same manner as the P-oxidation of saturated fatty acids, with successive cleavage of two-carbon units as acetyl-CoA However, after three cycles of P-oxidation, the chain-shortening process produces an acyl-CoA molecule that has a cis-3,4 double bond At this point, an ancillary enzyme, A3,A2enoyl-CoA isomerase, converts the A3,4-cisdouble bond to a A2, -truns double bond, thus providing a suitable substrate for enoyl-CoA hydratase A slightly different situation arises when the chain-shortening process produces an acyl-CoA molecule that has a cis-4,5 double bond.

Offshoring. In 2004, by far the hottest outsourcing topic fueling political debates at the local, state, and federal levels and at times recognized as a major issue by the presidential candidates has been offshore outsourcing or offshoring. Although offshoring has been around for several years, the seemingly recent predictions made by outsourcing analysts regarding the growth in the offshore outsourcing industry have been meet with astonishment and calls by the legislature to limit offshoring in an effort to keep jobs in the United States (particularly outsourcing efforts by federal and state government entities).7 Examples of some of the predictions include the following:

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Under these conditions, the acyl-CoA dehydrogenase step generates a A2~3-transl A4, -cis conjugated diunsaturated fatty acyl-CoA intermediate (Fig 10-7) At this point a second ancillary enzyme, NADPH-dependent 2,4-dienoyl-CoA reductase, transfers hydrogen atoms from NADPH to carbons 4 and 5, generating a A bms-enoyl-CoA A3,A2-Enoyl isomerase then converts A3~4-truns-enoyl-CoA A2~ -trans-enoy1-CoA to Note that by virtue of the carbon-carbon double bonds they contain, unsaturated fatty acids generate slightly less ATP than do saturated fatty acids For example, @-oxidationof oleic acid bypasses one acyl-CoA dehydrogenase step and thereby generates one less FADH2 molecule (and 15 fewer ATP) than its saturated counterpart, stearic acid 10435 Oxidation of Medium-Chain Fatty Acids Breast milk contains relatively large amounts of medium-chain (CX-C 12) fatty acids that provide nursing infants with substantial amounts of energy.

These shorter fatty acids are more soluble than are their more common C16-C20 counterparts and can enter the mitochondrion directly from the cytosol without need for the carnitine transporter system The C8-C 12 fatty acids are activated to their corresponding acyl-CoA derivatives within the mitochondrion and then undergo P-oxidation The initial oxidation reaction is catalyzed by medium-chain acyl-CoA dehydrogenase (MCAD) Because mediumchain fatty acids bypass both the carnitine transporter system and the @-oxidation enzymes specific to long-chain acyl-CoAs, TAG comprised of CX-C12 fatty acids are sometimes used as nutrient supplements to treat persons who cannot oxidize long-chain fatty acids due to genetic deficiencies of proteins involved in either the transport or oxidation of long-chain fatty acids 10436 Oxidation of Fatty Acids Containing an Odd Number Of Carbon Atoms Dietary lipids often contain a small amount of odd-carbon fatty acids such as 17:O.

Thus EC decay is favored for high Z nuclei. Of course, the decay energy must be greater than 1.02 MeV for b decay, a situation found mostly in low Z nuclei where the slope of the walls of the valley of b stability is large (see Fig. 2.8) and decay energies of .1.02 MeV occur. Electron capture decay produces a vacancy in the atomic electron shells and secondary processes that lead to lling that vacancy by the emission of X-rays and Auger electrons occur. These X-rays permit the detection of EC decays.

Odd-chain fatty acids are oxidized by the normal mitochondria1 P-oxidation process However, the last thiolytic cleavage step produces one molecule of acetylCoA and one molecule of propionyl-CoA from the methyl end of the fatty acid As indicated in 9, propionyl-CoA is gluconeogenic in a fasting person because it can be metabolized to glucose in the liver The incorporation of propionyl-CoA.

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The SLAX Popcorn bootable CD image is included on the CD that comes with this book. This live CD contains a well-stocked desktop Linux system. SLAX is described in 19. To run SLAX live, insert the CD into your PC s CD drive, reboot, and type the following from the boot prompt:

If you become a Linux administrator or power user, over time you will inevitably find yourself working on a computer from a remote login or where there is no desktop GUI available At some point while you are in that state, you will want to check an HTML file or a Web page To solve the problem, many Linux distributions include several text-based Web browsers With text-based Web browsers, any HTML file available from the Web, your local file system, or a computer where you re remotely logged in can be accessed from your shell There s no need to fire up your GUI or read pages of HTML markup if you just want to take a peek at the contents of a Web page.

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